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11 Pág. ; Maternal effects have been demonstrated to affect offspring performance in many organisms, and in plants, seeds are important mediators of these effects. Some woody plant species maintain long-lasting canopy seed banks as an adaptation to wildfires. Importantly, these seeds stored in serotinous cones are produced by the mother plant under varying ontogenetic and physiological conditions. ; This research was supported by the Spanish Government via the Ministry of Science, Innovation and Universities (MICIU) grants AGL2015-68274-C03-1-R and RTI2018-094691-B-C32 and by a predoctoral research fellowship from the Spanish MICIU (BES-2016-077347). ; Peer reviewed