Suchergebnisse

The interactions between bees that depend on floral oil for their larvae and flowers that offer oil involve an intricate mix of obligate and facultative mutualisms. Using recent phylogenies, new data on oil-offering Cucurbitaceae, and molecular-dating, we ask when and how often oil-offering flowers and oil-foraging bees evolved, and how frequently these traits were lost in the cause of evolution. Local phylogenies and an angiosperm-wide tree show that oil flowers evolved at least 28 times and that floral oil was lost at least 36–40 times. The oldest oil flower systems evolved shortly after the K/T boundary independently in American Malpighiaceae, tropical African Cucurbitaceae and Laurasian Lysimachia (Myrsinaceae); the ages of the South African oil flower/oil bee systems are less clear. Youngest oil flower clades include Calceolaria (Calceolariaceae), Iridaceae, Krameria (Krameriaceae) and numerous Orchidaceae, many just a few million years old. In bees, oil foraging evolved minimally seven times and dates back to at least 56 Ma (Ctenoplectra) and 53 Ma (Macropis). The co-occurrence of older and younger oil-offering clades in three of the four geographical regions (but not the Holarctic) implies that oil-foraging bees acquired additional oil hosts over evolutionary time. Such niche-broadening probably started with exploratory visits to flowers resembling oil hosts in scent or colour, as suggested by several cases of Muellerian or Batesian mimicry involving oil flowers.

Open Access article, published by EDP Sciences, under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. ; Context. The tenuous nitrogen (N2) atmosphere on Pluto undergoes strong seasonal effects due to high obliquity and orbital eccentricity, and has recently (July 2015) been observed by the New Horizons spacecraft. Aims. The main goals of this study are (i) to construct a well calibrated record of the seasonal evolution of surface pressure on Pluto and (ii) to constrain the structure of the lower atmosphere using a central flash observed in 2015. Methods. Eleven stellar occultations by Pluto observed between 2002 and 2016 are used to retrieve atmospheric profiles (density, pressure, temperature) between altitude levels of ∼5 and ∼380 km (i.e. pressures from ∼10 μbar to 10 nbar). Results. (i) Pressure has suffered a monotonic increase from 1988 to 2016, that is compared to a seasonal volatile transport model, from which tight constraints on a combination of albedo and emissivity of N2 ice are derived. (ii) A central flash observed on 2015 June 29 is consistent with New Horizons REX profiles, provided that (a) large diurnal temperature variations (not expected by current models) occur over Sputnik Planitia; and/or (b) hazes with tangential optical depth of ∼0.3 are present at 4-7 km altitude levels; and/or (c) the nominal REX density values are overestimated by an implausibly large factor of ∼20%; and/or (d) higher terrains block part of the flash in the Charon facing hemisphere.© E. Meza et al. 2019 ; The work leading to these results has received funding from the European Research Council under the European Community's H2020 2014-2020 ERC Grant Agreement no 669416 >Lucky Star>. E.M. thanks support from Concytec-Fondecyt-PE and GA, FC-UNI for providing support during the 2012 July 18 occultation. B.S. thanks S. Para for partly supporting this research though a donation, J. P. Beaulieu for helping us accessing to the Hobart Observatory facilities and B. Warner, B. L. Gary, C. Erickson, H. Reitsema, L. Albert, P. J. Merritt, T. Hall, W. J. Romanishin, Y. J. Choi for providing data during the 2007 March 18 occultation. M.A. thanks CNPq (Grants 427700/2018-3, 310683/2017-3 and 473002/2013-2) and FAPERJ (Grant E-26/111.488/2013). J.L.O. thanks support from grant AYA2017-89637-R. P.S.S. acknowledges financial support from the European Union's Horizon 2020 Research and Innovation Programme, under Grant Agreement no 687378, as part of the project > Small Bodies Near and Far> (SBNAF). J.L.O., R.D., P.S.S. and N.M. acknowledge financial support from the State Agency for Research of the Spanish MCIU through the > Center of Excellence Severo Ochoa> award for the Instituto de Astrofisica de Andalucia (SEV-2017-0709). F.B.R. acknowledges CNPq support process 309578/2017-5. G.B.R. thanks support from the grant CAPES-FAPERJ/PAPDRJ (E26/203.173/2016). J.I.B.C. acknowledges CNPq grant 308150/2016-3. R.V.M. thanks the grants: CNPq-304544/2017-5, 401903/2016-8, and Faperj: PAPDRJ-45/2013 and E-26/203.026/2015. B.M. thanks the CAPES/Cofecub-394/2016-05 grant and CAPES/Brazil -Finance Code 001. B.M. and A.R.G.J. were financed in part by the Coordenacao de Aperfeicoamento de Pessoal de Nivel Superior -Brasil (CAPES) -Finance Code 001. TRAPPIST-North is a project funded by the University of Liege, in collaboration with Cadi Ayyad University of Marrakech (Morocco). TRAPPIST-South is a project funded by the Belgian Fonds (National) de la Recherche Scientifique (F.R.S.-FNRS) under grant FRFC 2.5.594.09. F, with the participation of the Swiss National Science Foundation (FNS/SNSF). VSD, SPL, TRM and ULTRACAM are all supported by the STFC. K.G. acknowledges help from the team of Archenhold-Observatory, Berlin, and A. R. thanks G. Roman (Granada) for help during the observation of the 2016 July 19 occultation. A.J.C.T. acknowledges support from the Spanish Ministry Project AYA2015-71718-R (including EU funds). ; Peer Reviewed