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Copyright: © 2013 Tenesa et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. Funding: Genotyping of the ABC1936, LBC1921, and LBC1936 cohorts and the analyses conducted here were supported by the UK's Biotechnology and Biological Sciences Research Council (BBSRC). Phenotype collection in the Lothian Birth Cohort 1921 was supported by the BBSRC, The Royal Society, and The Chief Scientist Office of the Scottish Government. Phenotype collection in the Lothian Birth Cohort 1936 was supported by Research Into Ageing (continues as part of Age UK's The Disconnected Mind project). Phenotype collection in the Aberdeen Birth Cohort 1936 was supported by the BBSRC, the Wellcome Trust, and the Alzheimer's Research Trust. SJR, AR and AT are funded by the BBSRC through the Roslin Institute's strategic programme grant and project grant BB/K000195/1. The Brain data was provided by the Division of Aging Biology and the Division of Geriatrics and Clinical Gerontology (NIA) through the NIH GWAS Data Repository (dbGaP Accession Number: phs000249.v1.p1) and funded as part of the Intramural Research Program, NIA. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. ; Peer reviewed ; Publisher PDF